Plant tissues are of two types on the basis of their dividing capacity :
1. Meristematic tissue (growing tissue)
2. Permanent tissue
- These are living tissues which are composed of immature cells that are capable of division throughout life.
- These tissues are found in growing regions of plants.
- Cells have thin cell wall.
- Cells contain dense cytoplasm and do not have vacuoles.
- Cells contain prominent and large nucleus.
- Cells are metabolically highly active, so store food is absent.
- Cells are compactly arranged because they do not have intercellular spaces.
- Function : Meristematic tissue is responsible for the growth in length and width(girth) of plant body.
➢ Classification of Meristematic Tissue
On the basis of their location meristematic tissues are of three types:
1. Apical Meristem
- It is present at the growing tips of stems and roots.
- They are responsible for increase in the length of plant organs.
- Apical meristem is absent in Algae and Fungi.
- Apical and intercalary meristem are responsible for primary growth of plant (growth in length).
Meristematic tissues are of three types on the basis of origin-
Promeristem : These are embryonic meristem which give rise to primary meristem.
Primary Meristem: These are always in active state of division and give rise to primary permanent tissue.
Secondary Meristem: These are developed from primary permanent tissue and give rise to secondary permanent tissues.
Do You Know
Shoot apical meristem is terminal in position.
Root apical meristem is subterminal in position due to the presence of root cap.
Apical meristem has two regions at embryonic stage.
First - Promeristem Second - Eumeristem
Eumeristem is divided into three regions
1. Protoderm 2. Procambium 3. Ground meristem
2. Intercalary Meristem
- It is the part of apical meristem which is left behind during growth period. They are short lived and convert into permanent tissue.
- These are present at the base of leaf or internode. Intercalary meristem may be present either at the node as in grasses, bamboo and mints or the base of Pinus leaves.
- They are responsible for the growth in length of plant organs.
3. Lateral Meristem
- It lies on the lateral sides of stem and root or occurs along the sides of longitudinal axis of the plant.It helps in increasing the diameter (girth or width) of plant. Hence helps in secondary growth.
Lateral meristem are both primary and secondary in origine.
Primary lateral meristem e.g marginal meristem of leaf and Intra fascicular cambium.
Secondary lateral meristem e.g. Inter fascicular cambium and cork cambium. Lateral meristem is present in only dicot plants.
- They are formed by division and differentiation of meristematic tissue.
- They are composed of those cells which have lost the power of division (temporarily or permanent) and attain a permanent shape, size and function. Cells may be living or dead.
- Cells may be oval, rounded, polygonal or elongated.
- Permanent tissues are of two types :
(i) Simple permanent tissue
(ii) Compound or Complex Permanent Tissue
➢ Simple Permanent Tissue
- These tissues are made up of similar types of cells, that perform a common function.
- They are protective and supportive in nature.
- Simple tissues are of three types :
- It is a living and basic packing tissue which consists of relatively unspecialised cells.
- Cells of these tissues have thin cell wall which is made up of cellulose.
- Cells of these tissues have dense cytoplasm with small nucleus and large vacoule.
- They are usually loosely packed because intercellular spaces are present between cells.
- Functions : Storage of food and provide support to the plant.
Do You Know
Parenchyma is the first evolved permanent tissue which is present in all soft parts of plant (therefore called as universal tissue).The body of bryophytes is made up of only parenchyma tissue. In dorisventral leaf of dicot plant, two types of parenchyma tissues are present:
(i) spongy tissue (ii) palisade tissue
Parenchyma provides targidity to cells.
Parenchyma cells which store rasin tanin gum and oil are also so called idioblast.
Prosenchyma : Parenchymatous cells become long and taper at both the ends.e.g. It is found in pericycle of root.
Stellate parenchyma found in the leaf bases of banana, leaf bases performs the function of stem.
Differentiation : The developement process in which cells take up a permanent shape, size and perform a specific function.
Modification of parenchyma :
- Such type of parenchyma in which abundant quantity of chloroplasts are found. (contains chloroplasts)
- They are present in mesophyll of leaves.
- Function : Synthesis of food (Photosynthesis)
- Parenchyma is made up of rounded cells which surrounds the large air cavities. It is found in aquatic plants or hydrophytes.
- Example : petiole of water hyacinth.
- Function : It provides buoyancy to the aquatic plants to help them float.
2. Collenchyma (Flexible tissue)
- Cells of this tissue are living, elongated or vary in structure. Cells of this tissue are irregularly thickend at the corners due to the deposition of pectin.
- It is present below the epidermis of leaf and harbaceous stem in the form of hypodermis.
- Intercellular spaces are very little or absent between cells of this tissue.
- Collenchyma are usually absent in monocots and in roots.
- Cells of collenchyma contain few chloroplast.
- Functions :
(i) It provides mechanical support (tensile strength) and elasticity.
(ii) It protects the cracking of lamina margin due to action of wind.
(iii) It provides flexibility to plant.
(iv) It allows easy bending in various parts of plant (leaf and stem) without breaking.
- Sclerenchyma cells are dead cells and they are devoid to protoplasm.
- The walls of cells of sclerenchyma are greatly thickened with deposition of lignin. Such cell walls are called lignified.
- Due to excessive thickening of the wall of a sclerenchymatous cell, its cavity or lumen becomes nearly absent. The cells of sclerenchyma are closely packed without intercellular spaces.
- They are found in stems (around the vascular bundle), roots, veins of leves, hard coverings of seed and nuts.
- Function : It is the main mechanical tissue which provides mechanical support.
Sclerenchymatous cells are of two types in structure :
(a) Sclerenchyma fibres
- They are highly elongated (1 mm to 550 mm in length), narrow and spindle shaped with pointed or oblique end walls.
- The fibres are closely packed without intercellular spaces.
- The walls of fibres are often so thick that the cell cavity or lumen is nearly absent. Often oblique thin areas are found in their walls. These are called pits.
- Functions :
(i) Sclerenchyma fibres constitute the major mechanical tissue of the plants and are abundantly found in plants.
(ii) Commercial fibres obtained from plants (e.g. Jute, Flax, Hemp, Husk of coconut etc.) usually are sclerenchymatous fibres.
Lignin is a complex polymer which acts as a cement and hardens cell wall. Lignin makes the cell wall impermeable so important substances are unable to pass through it. As a result, cells that are heavily lignified do not have living content (protoplasm).
(b) Sclereids (grit or stone cells)
- They are highly thickened and irregularly shaped dead cells.
- They are found in various parts of the plant such as cortex, pith, phloem, hard seeds etc.
- The grit of pulp of some fruits (such as guava, apple, pear etc.) is due to the presence of sclereids in it.
- Functions :
(i) The main function of sclerenchyma is to provide mechanical support to the plants.
(ii) Sclereids provide strength to seed covering and grittiness to the pulp of many fruits.
Differences Between Fibres and Sclereids
➢ Compound or Complex Permanent Tissue
- The complex tissues consist of more than one type of cells. All these cell co-ordinate to perform a common function.
- Complex tissues transport water, mineral salts (nutrients) and food material to various parts of plant body.
- Complex tissues are of the following two types :
(i) Xylem or wood
(ii) Phloem or bast
1. Xylem or Wood
Xylem is made up of four types of cells :
- Tracheids are elongated cells with tapering ends.
- They also conduct water. Since tracheids do not have open ends like the vessels, so the water has to pass from cell to cell via the pits.
(b) Vessels or Tracheae
- Very long tube like structures formed by a row of cells placed end to end.
- The transverse walls between the vessels are completely dissolved to form continuous channels or water-pipes.
- Functions :
(i) Tracheids and vessels help in long distance conduction of water and minerals upward from the root system to various parts of plant.
(ii) Tracheids and vessels provides mechanical support.
(c) Xylem Fibre
- These are dead and lignified sclerenchymatous cells which are mainly supportive in function.
(d) Xylem Parenchyma
- It is formed of living parenchymatous cells which helps in storage of food and lateral conduction of water and minerals.
Do You Know
Xylem and phloem are both conducting tissues and also known as vascular tissues (conducting tissue), together both of them constitute vascular bundles.
Vessels and tracheids have 5 type of lignification.
Annular, spiral, reticulate, scalariform and pitted.
Vessels are only found in xylem of angiosperm but exceptionally present in gymnosperm (Ephedra, Gnetum and Welwitschia). Vessels are example of dead syncyte.
- No lignin is laid down where plasmodesmata were present in the original cell walls.
- These non-lignified areas are known as pits and they allow water to pass sideways between one xylem vessel and the next.
- As vessels and tracheids of xylem have the lignified cell walls, so this simply mean that these cells are hollow and there are no cell contents to restrict the flow of water.
- Tracheids & vessels are collectively called as hadrome and the hadrome as main conducting elements in xylem.
- The annual rings present in the trunk of a tree are xylem rings. By counting the number of annual rings we can determine the age of a tree (Dendrocronology).
It is also made up of four types of cells:
(a) Sieve tubes
- Sieve tubes are slender, tube-like structures composed of elongated thin-walled cells, placed end to end.
- Their end walls are perforated by numerous pores and are called sieve plates. Walls of sieve tubes are perforated.
- The nucleus of each sieve cell degenerates at maturity, however, cytoplasm persists in the mature cell. Thus, nuclei are absent in mature sieve tube elements.
- The cytoplasm of one sieve tube element is continuous with those of the sieve elements above and below by cytoplasmic connections passing through the pores of the sieve plate.
Although sieve tube elements do not have nuclei, but they still remain living. It is so because they are dependent on adjacent companion cells which develop from the same original meristematic cell. The two cells together for a functional unit.
Leptome : Main part of phloem involved in conduction of food, which is sieve tube. companion cells are present only in phloem of angiospermic plants.
(b) Companion cells
- These are associated with sieve tubes. These are smaller cells having dense cytoplasm and prominent nucleus.
- The companion cells help the sieve tubes in the conduction of food material. Sieve cells & companion cells are so called sister cell because they originate from single mother cell.
(c) Phloem parenchyma
- These are living and thin walled cells. It is also known as bast parenchyma.
- It helps in conduction of food in radial direction.
- It store various materials e.g. Resin, Latex, Mucilage.
(d) Phloem fibres
- These are dead and sclerenchymatous cells. Phloem or bast fibres of some plants are ource of commercial fibres e.g. Jute, Hemp, Flex.
- They provide mechanical support to the conducting elements.
Function of Phloem : Phloem transport photosynthetically prepared food materials from the leaves to the storage organs and latter from storage organs to the growing regions of the plant body.
- These tissues are primarily protective in function.
- They are of two types :
It is the outermost layer of all organs of plant body which is formed from parenchymal cells. It protects the internal tissue from mechanical injuries and entry of germs.
- The outer wall of epidermis of aerial parts of plant is deposited with a fatty substance, called cutin which form a water proof layer called cuticle.
- It checks the loss of water by transpiration.
- Lower epidermis of dicot leaves have large number of microscopic aperture called stomata.
- Each stomatum is an elliptical apperture bounded by two kidney shaped guard cells which regulate opening and closing stomatum.
- Guard cells are kidney shaped in dicot and dumb-bell shaped in monocot.
- Stomata helps in exchange of gases.
- It helps in loss of water vapours called transpiration, develops a force called transpiration pull,which helps in absorbtion of water by the roots.
➢ Root hairs
- Epidermis of roots (epiblema) have root hairs which greatly increase their surface area for absorption of water and minerals.
2. Cork or Phellem
- Cork is the peripheral tissue of old stems and roots of woody trees and is formed due to activity of cork cambium or phellogen (secondary lateral meristem).
- Cork cambium produces off new cell on its both sides, thus, forming cork (phellem) on the outer side and the Secondary cortex or phelloderm on the inner side.
- It is made up of dead cells with thick wall but no intercellular spaces.
- The walls of cork cells are heavily thickened by the deposition of an organic substance (a fatty substance), called suberin.
- Suberin makes these cell impermeable to water and gases and it also helps in conservation of water in the trees.
➢ Commercial Importance of Cork
- Cork is light and highly compressible which does not catch fire easily.
- Cork is used in the making of a variety of sports goods such as cricket balls, table tennis, shuttle-cocks, wooden paddles etc.
- Cork is protective in function.
- Cork prevents desiccation (by preventing loss of water from plant.)
- Cork prevents infection and mechanical injury.
- Lenticels (pores) present in the cork provide aeration to the inner tissues.
Dedifferentiation : In this process the specialized cells regain the division power and become meristematic e.g. vascular cambium
The cork cells do not contain protoplasm but are filled with resin or tannins. In case of onion bulb too, in the skin of onion the cell walls become thick and water proof due to addition of suberin.
Cork and bark : are not the same structures. While cork includes outer products of cork cambium, the bark includes the outer products of cambium such as secondary phloem and also cork cambium and cork.
Commercial cork is obtained from the stem surface of Quercus suber.