Page 1
Institute of Lifelong Learning, University of Delhi
Lesson Prepared Under MHRD project “National Mission on
Education Through ICT”
Discipline: Botany
Paper: Plant Anatomy
National Coordinator: Prof. S.C. Bhatla, University of Delhi
Lesson: Morphology of Root, Stem and Leaf
Lesson Developer: Dr. Pooja Gokhale Sinha and Dr. Ajit
Kumar Kavathekar
Department/College: Sri Venkateswara College, University
of Delhi
Lesson Reviewer: Prof. S.C. Bhatla
Department/College: Botany Department
Language Editor: Dr. Arun Kumar Maurya
Department/College: Botany Department, Dyal Singh
College,
University of Delhi
Lesson Editor: Dr. Rama Sisodia, Fellow in Botany ILLL
Page 2
Institute of Lifelong Learning, University of Delhi
Lesson Prepared Under MHRD project “National Mission on
Education Through ICT”
Discipline: Botany
Paper: Plant Anatomy
National Coordinator: Prof. S.C. Bhatla, University of Delhi
Lesson: Morphology of Root, Stem and Leaf
Lesson Developer: Dr. Pooja Gokhale Sinha and Dr. Ajit
Kumar Kavathekar
Department/College: Sri Venkateswara College, University
of Delhi
Lesson Reviewer: Prof. S.C. Bhatla
Department/College: Botany Department
Language Editor: Dr. Arun Kumar Maurya
Department/College: Botany Department, Dyal Singh
College,
University of Delhi
Lesson Editor: Dr. Rama Sisodia, Fellow in Botany ILLL
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 1
Table of Contents
Chapter: Plant Morphology: Morphology of Root, Stem and Leaf
? Introduction
? Plant Body Organization
? Organs
? Tissues
? Root System and their modifications
? Shoot System and their modifications
? Summary
? Exercise/ Practice
? Glossary
? References/ Bibliography/ Further Reading
Page 3
Institute of Lifelong Learning, University of Delhi
Lesson Prepared Under MHRD project “National Mission on
Education Through ICT”
Discipline: Botany
Paper: Plant Anatomy
National Coordinator: Prof. S.C. Bhatla, University of Delhi
Lesson: Morphology of Root, Stem and Leaf
Lesson Developer: Dr. Pooja Gokhale Sinha and Dr. Ajit
Kumar Kavathekar
Department/College: Sri Venkateswara College, University
of Delhi
Lesson Reviewer: Prof. S.C. Bhatla
Department/College: Botany Department
Language Editor: Dr. Arun Kumar Maurya
Department/College: Botany Department, Dyal Singh
College,
University of Delhi
Lesson Editor: Dr. Rama Sisodia, Fellow in Botany ILLL
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 1
Table of Contents
Chapter: Plant Morphology: Morphology of Root, Stem and Leaf
? Introduction
? Plant Body Organization
? Organs
? Tissues
? Root System and their modifications
? Shoot System and their modifications
? Summary
? Exercise/ Practice
? Glossary
? References/ Bibliography/ Further Reading
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 2
Learning Objects
After studying this unit, you should be able to:
? Understand and define the term morphology.
? Recognize the variability in forms.
? Correlate form to the functions they perform.
? Distinguish between root and shoot system.
? Comprehend that organs are made up of different kinds of tissues.
? Visualize a tissue, an organ or a plant as a dynamic entity sensitive to its
surrounding environment.
Page 4
Institute of Lifelong Learning, University of Delhi
Lesson Prepared Under MHRD project “National Mission on
Education Through ICT”
Discipline: Botany
Paper: Plant Anatomy
National Coordinator: Prof. S.C. Bhatla, University of Delhi
Lesson: Morphology of Root, Stem and Leaf
Lesson Developer: Dr. Pooja Gokhale Sinha and Dr. Ajit
Kumar Kavathekar
Department/College: Sri Venkateswara College, University
of Delhi
Lesson Reviewer: Prof. S.C. Bhatla
Department/College: Botany Department
Language Editor: Dr. Arun Kumar Maurya
Department/College: Botany Department, Dyal Singh
College,
University of Delhi
Lesson Editor: Dr. Rama Sisodia, Fellow in Botany ILLL
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 1
Table of Contents
Chapter: Plant Morphology: Morphology of Root, Stem and Leaf
? Introduction
? Plant Body Organization
? Organs
? Tissues
? Root System and their modifications
? Shoot System and their modifications
? Summary
? Exercise/ Practice
? Glossary
? References/ Bibliography/ Further Reading
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 2
Learning Objects
After studying this unit, you should be able to:
? Understand and define the term morphology.
? Recognize the variability in forms.
? Correlate form to the functions they perform.
? Distinguish between root and shoot system.
? Comprehend that organs are made up of different kinds of tissues.
? Visualize a tissue, an organ or a plant as a dynamic entity sensitive to its
surrounding environment.
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 3
Introduction
In “considering the distinctive characters of plants and their nature generally one must take
into account their parts, their qualities, the ways in which their life originate and the course
which it follows in each case…..”
Above is the translation by Sir Arthur Hort (1949) of the opening sentence of the famous
book “Enquiry in Plants” by Theophrastus, written more than 300 B.C. ago (Sporne, 1965).
Such an expression of plant body structure by Theophrastus has probably laid the basic
foundation of the field morphology. Eminent botanist A. Arber (1946) considered such
comprehensive description remarkable as it encompasses modern scientific fields, such as
anatomy, physiology, biochemistry, reproduction and life histories. In essence, the form or
the look is a combined manifestation of all of the above. Literally the word „Morphology? is
ancient Greek in origin. However, term morphology in biological context was first used by a
German poet and Philosopher J. W. Goethe in 1790. It means “study of form”. Simply put,
plant morphology can be defined as the study of physical form and external features of a
plant.
What does the word “form” mean? It is simply defines how any object looks. Crudely, it
reflects its shape. For an example, if you were asked to draw a picture of moon, how shall
you draw? In all probability your drawings shall reflect a circle or a crescent, but never a
rectangle or a triangle. Similarly if you were to draw a pencil, you invariably make a rod-like
shape with a pointed end but in case of banana fruit, sketch shall depict a curve. All of us,
thus perceive an object, be it living or non-living, to a particular “form”. This form is what
one can call or describe as the “morphology” of that object. Can we say then that there are
or could be as many forms as are the number of objects?
Even early man, probably, used the „form? to distinguish edible and poisonous plants. How
could they do it? Morphology must have been the most significant basis of distinction. The
field of morphology must have been developing over a long period of time even before man
Page 5
Institute of Lifelong Learning, University of Delhi
Lesson Prepared Under MHRD project “National Mission on
Education Through ICT”
Discipline: Botany
Paper: Plant Anatomy
National Coordinator: Prof. S.C. Bhatla, University of Delhi
Lesson: Morphology of Root, Stem and Leaf
Lesson Developer: Dr. Pooja Gokhale Sinha and Dr. Ajit
Kumar Kavathekar
Department/College: Sri Venkateswara College, University
of Delhi
Lesson Reviewer: Prof. S.C. Bhatla
Department/College: Botany Department
Language Editor: Dr. Arun Kumar Maurya
Department/College: Botany Department, Dyal Singh
College,
University of Delhi
Lesson Editor: Dr. Rama Sisodia, Fellow in Botany ILLL
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 1
Table of Contents
Chapter: Plant Morphology: Morphology of Root, Stem and Leaf
? Introduction
? Plant Body Organization
? Organs
? Tissues
? Root System and their modifications
? Shoot System and their modifications
? Summary
? Exercise/ Practice
? Glossary
? References/ Bibliography/ Further Reading
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 2
Learning Objects
After studying this unit, you should be able to:
? Understand and define the term morphology.
? Recognize the variability in forms.
? Correlate form to the functions they perform.
? Distinguish between root and shoot system.
? Comprehend that organs are made up of different kinds of tissues.
? Visualize a tissue, an organ or a plant as a dynamic entity sensitive to its
surrounding environment.
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 3
Introduction
In “considering the distinctive characters of plants and their nature generally one must take
into account their parts, their qualities, the ways in which their life originate and the course
which it follows in each case…..”
Above is the translation by Sir Arthur Hort (1949) of the opening sentence of the famous
book “Enquiry in Plants” by Theophrastus, written more than 300 B.C. ago (Sporne, 1965).
Such an expression of plant body structure by Theophrastus has probably laid the basic
foundation of the field morphology. Eminent botanist A. Arber (1946) considered such
comprehensive description remarkable as it encompasses modern scientific fields, such as
anatomy, physiology, biochemistry, reproduction and life histories. In essence, the form or
the look is a combined manifestation of all of the above. Literally the word „Morphology? is
ancient Greek in origin. However, term morphology in biological context was first used by a
German poet and Philosopher J. W. Goethe in 1790. It means “study of form”. Simply put,
plant morphology can be defined as the study of physical form and external features of a
plant.
What does the word “form” mean? It is simply defines how any object looks. Crudely, it
reflects its shape. For an example, if you were asked to draw a picture of moon, how shall
you draw? In all probability your drawings shall reflect a circle or a crescent, but never a
rectangle or a triangle. Similarly if you were to draw a pencil, you invariably make a rod-like
shape with a pointed end but in case of banana fruit, sketch shall depict a curve. All of us,
thus perceive an object, be it living or non-living, to a particular “form”. This form is what
one can call or describe as the “morphology” of that object. Can we say then that there are
or could be as many forms as are the number of objects?
Even early man, probably, used the „form? to distinguish edible and poisonous plants. How
could they do it? Morphology must have been the most significant basis of distinction. The
field of morphology must have been developing over a long period of time even before man
Plant Morphology: Morphology of root, stem and leaf
Institute of Lifelong Learning, University of Delhi 4
became civilized. We may also realize that no two natural forms are exactly same or alike.
At the same time some forms become characteristics of certain objects. Can you make a list
of such variations and conformations in the plants of your neighborhood?
The current chapter shall focus on the study of form, structure and organization of
angiosperms. Angiosperms or flowering plants are the most diverse group of land plants
that produce seeds in an enclosure of fruit wall. They differ from gymnosperms as they
possess vessels for efficient conduction of water, flowers with different whorls, ovules
enclosed in carpels, different mechanism of pollination and seed dispersal and double
fertilization.
Plant Body Organization
Let us now discuss the body organization of a flowering plant. How do we choose a plant or
a particular group of plants for our study? We generally use the phrase- “study of the
morphology of a typical flowering plant”. Angiosperms could be annual (pulses), biennial
(cabbage) or perennial (trees). It may be referred to as herb (potato, mustard); shrub
(Dodonia, Murraya, Bougainvillea, Rose) or tree (Banyan, Mango) (Figure 1). An angiosperm
could be as small as Wolffia (2 mm across) or as tall as 100 feet Eucalyptus. It could inhabit
diverse phytogeographical regions, viz. warm tropics (banana, papaya, orchids); temperate
mountains Rhododendron; deserts (cacti); water bodies (water hyacinth), estuarine zones
(mangroves); crevices and rocks (Phlox, Alyssum, Primula); climbers, epiphytes (orchids),
and so on (Figure 2). Above demarcation of categories to which a plant is referred thus
depends on the parameter under consideration.
You may well wonder are we going to study morphology of all these plants or groups of
plant? If yes, how? If no, why? There are around 13,000 genera and 2,40,000 plant species
inhabiting our planet (Takhtajan, 1980). It must be emphasized here that whatever be the
size, age, the number or kinds of organs, tissues or cells a plant possesses at a given time
or spread over a certain period of time, which or how many of these can be considered as
an ideal to study body organization? For simplicity, we choose a model set of body
organizational set-up that most of the plants possess. This set up is termed as a typical.
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