Population Attributes
- In nature, most individuals of a species are found living in groups within a specific geographical area. These individuals share or compete for similar resources and have the potential to interbreed, thus forming a population. While interbreeding typically refers to sexual reproduction, groups of individuals resulting from asexual reproduction are also considered populations for ecological studies. Examples of populations include all the cormorants in a wetland, rats in an abandoned dwelling, teakwood trees in a forest, bacteria in a culture plate, and lotus plants in a pond.
- Population ecology is crucial because it connects ecology with population genetics and evolution. Although individual organisms must adapt to environmental changes, natural selection operates at the population level to evolve desired traits. Populations have specific attributes that individuals do not. For instance, while an individual may experience births and deaths, a population has birth rates and death rates, which are expressed as per capita rates. These rates indicate changes in numbers (increase or decrease) concerning the members of the population.
(i) Birth and Death Rates
Birth and death rates are crucial attributes of a population. While an individual organism may experience births and deaths, a population is characterized by birth rates and death rates, which are expressed per capita. These rates reflect the change in numbers (increase or decrease) concerning the members of the population.
Example:
- Birth Rate: Imagine a pond with 20 lotus plants last year. If 8 new plants are added through reproduction, the current population becomes 28. The birth rate is calculated as 8/20 = 0.4 offspring per lotus per year.
- Death Rate: In a laboratory population of 40 fruit flies, if 4 individuals die during a specified time interval (e.g., a week), the death rate is 4/40 = 0.1 individuals per fruit fly per week.
(ii) Sex Ratio
A population's sex ratio indicates the proportion of males and females within it. For instance, if 60 percent of the population are females and 40 percent are males, this distribution reflects the sex ratio of the population.
(iii) Age Distribution and Age Pyramid
Representation of age pyramids for human population
- Age Distribution: The age distribution of a population, represented as the percentage of individuals of different ages or age groups, provides insights into the population's structure.
- Age Pyramid: A graphical representation of age distribution is called an age pyramid. For human populations, age pyramids typically display the age distribution of males and females. The shape of the pyramid indicates whether the population is growing, stable, or declining.
(iv) Population Size and Population Density
- Population Size: The total number of individuals in a population at a given time.
- Population Density (N): Population size can be expressed as population density, which may not always be measured in numbers. While total number is often the most appropriate measure, there are instances where it is meaningless or challenging to determine.
(v) Relative Measures and Indirect Estimation
- In some cases, relative measures of population size are sufficient. For instance, the number of fish caught per trap can indicate the total population density in a lake.
- Estimating population sizes indirectly is common. For example, tiger censuses in national parks and reserves often rely on pug marks and fecal pellets rather than direct counting.
Understanding these population attributes is essential for studying ecological processes, such as competition with other species, the impact of predators, or the effects of pesticide applications, all evaluated in terms of changes in population size.
Question for Population Attributes & Growth
Try yourself:
Which of the following population attributes is represented by the proportion of males and females within a population?Explanation
- The sex ratio of a population indicates the proportion of males and females within it. It is an important attribute used to understand the demographic structure of a population.
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Population Growth
Population growth refers to the change in the number of individuals in a population over time. It is influenced by factors such as birth rates, death rates, immigration, and emigration. Understanding these factors helps us analyze whether a population is increasing, decreasing, or stable.
Key Processes in Population Density Changes
Population density varies over time due to factors like food availability, predation, and weather.Changes in density indicate whether a population is thriving or declining.
Four basic processes affect population density:
- Natality: Number of births in a given period.
- Mortality: Number of deaths in a given period.
- Immigration: Individuals moving into a habitat from elsewhere.
- Emigration: Individuals leaving a habitat.
Population Density Equation
Population Density Key Processes
So, if N is the population density at time t, then its density at time t +1 is
Nt+1 = Nt + [(B + I) – (D + E)]
- Population density increases when births plus immigrants (B + I) exceed deaths plus emigrants (D + E).
- Under normal conditions, births and deaths are the main factors influencing population density.
- Immigration and emigration play a bigger role in specific situations, such as when a new habitat is being colonized.
Growth Models
Understanding how populations grow over time can help us learn about controlling population growth.
There are two main models to explain how populations grow:
(i) Exponential Growth
Exponential growth occurs when a population increases rapidly over time without any limits on resources like food and space.
Population growth curve a when responses are not limiting the growth, plot is exponential, b when responses are limiting the growth, plot is logistic, K is carrying capacity
Key Equation: The change in population size over time can be expressed with the formula:
dN/dt = (b - d) × N where:
dN/dt: Change in population size over time
b: Birth rate
d: Death rate
N: Population size
Intrinsic Rate of Natural Increase (r): This is a crucial factor in understanding population growth. It is calculated as r = b - d. For example, the Norway rat has an r value of 0.015, the flour beetle 0.12, and in 1981, India's human population had an r value of 0.0205.
J-shaped Curve: When plotted on a graph, exponential growth creates a J-shaped curve, indicating rapid population increase over time.
Integral Form: The integral form of the exponential growth equation is:
Nt = N0ert
where:Nt: Population density after time t
N0: Population density at time zero
r: Intrinsic rate of natural increase
e: Base of natural logarithms (approximately 2.71828)
Anecdote: To illustrate exponential growth, consider the story of a king and a minister playing chess. The minister bets on receiving wheat grains placed on the chessboard in a doubling pattern. This story highlights how quickly populations can grow when resources are unlimited, similar to how a tiny organism like Paramecium could multiply rapidly under ideal conditions.
Question for Population Attributes & Growth
Try yourself:
Which of the following factors does NOT influence population growth?Explanation
- Birth rates, immigration, and emigration are factors that influence population growth by affecting the number of individuals in a population.
- Weather, on the other hand, may indirectly impact population growth by influencing food availability and habitat suitability, but it is not a direct factor in determining population growth.
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(ii) Logistic Growth
- No population in nature has unlimited resources for exponential growth, leading to competition for limited resources.
- The ‘fittest’individuals survive and reproduce.
- Carrying Capacity (K): A habitat can support only a maximum number of individuals, called the carrying capacity (K).
- Phases of Growth: Initial lag phase, followed by phases of acceleration and deceleration, eventually reaching an asymptote when the population density hits K.
- Sigmoid Curve: A plot of population (N) against time (t) results in a sigmoid curve, indicative of logistic growth.
- Verhulst-Pearl Logistic Growth: Described by the equation:
- Where: N = Population density at time t,r = Intrinsic rate of natural increase,K = Carrying capacity.
- Logistic growth is considered more realistic because resources for growth are finite and become limiting over time.
- Government Census Data: Gather and plot population figures for India over the last 100 years to observe the evident growth pattern.
Life History Variation
- Populations evolve to maximize their reproductive fitness(high r value) in their specific habitat.
- Organisms evolve efficient reproductive strategies under particular selection pressures.
- Some organisms breed only once in their lifetime (e.g., Pacific salmon, bamboo), while others breed multiple times (e.g., most birds and mammals).
- Some produce a large number of small offspring (e.g., oysters, pelagic fishes), while others produce a small number of large offspring (e.g., birds, mammals).
- The desirability of breeding strategies for maximizing fitness depends on the constraints imposed by abiotic and biotic components of the habitat.
- The evolution of life history traits in different species is a significant area of research forecologists.
Question for Population Attributes & Growth
Try yourself:
Which of the following is a characteristic of logistic growth in a population?Explanation
- Logistic growth in a population is characterized by phases of acceleration and deceleration as the population approaches its carrying capacity.
- This pattern is more realistic as resources are finite and become limiting over time, leading to competition among individuals for survival and reproduction.
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