Gymnosperm
1. Term Gymnosperm was given by Theophrastus.
2. Study of Gymnosperm known as Gymnospermology.
3. Gymnosperm & Angiosperm are collectively included under spermatophyta i.e. seed-bearing plants.
4. Gymnosperms are naked seeded plant i.e. no fruit formation takes place in these plant. i.e. in gymnosperm embryo & seed formation takes place but no fruit formation occur.
5. Gymnosperms are very limited in distribution. They are mainly found in cold regions. In India, Gymnosperms are found on Himalayan mountains. They occur on slopes of the mountain in the cold region, therefore, gymnosperms are xerophyte.
6. All gymnosperm are vascular plants. Therefore vascular tissue present i.e. xylem & phloem. Xylem lack vessels & phloem lacks companion cells.
Note :
7. Most of the gymnosperms are arborescent (woody and tree habit) - but some are present as shrub. eg. Ephedra Some Gymnosperm are liana or woody climbers. eg. Gnetumula
Life Cycle Of Gymnosperm -
1. In Gymnosperms main plant body is sporophyte (diploid). All Gymnosperm are dioecious. i.e. male & female plants are separate, but exceptionally Pinus is monoecious.
2. All Gymnosperms are heterosporus. At the time of reproduction two types of spores are formed.
3. These two types of spores are formed in different sporangia.
4. Both types of sporangia are formed on different sporophylls.
5. Both types of sporophylls are found in groups & form male cone (strobilus) & female cone.
Carpel of Gymnosperm -
Carpels of Angiosperm -
6. Meiosis takes place in cells of microsporangium & megasporangium and form microspore & megaspore respectively.
7. In Gymnosperm and angiosperm germination of spores is endosporic i.e. germination of spore takes place with in sporangia.
8. Male gametophyte is formed after germination of microspore. Male gametophyte is also known as pollen grain.
Male gametophyte forms male gamete.
9. In lower gymnosperm, male gametes are motile & multiciliate, but in higher gymnosperm and angiosperm male gametes are non-motile due to the absence of cilia and flagella. They (higher gymnosperm) do not require water for fertilization.
10. Female gametophyte is formed after germination of megaspore.
Female gametophyte of gymnosperm contains two type of structures
11. Pollen grains (Male gametophyte) reach at the micropyle of ovule by wind pollination, called as onemophilly ovule by wind pollination.
Fertilization
Two types of fertilization take place in gymnosperms
(1) Zoodio - siphonogamy & This type of fertilization occurs in lower gymnosperms. Male gametes are motile and transferred to female gamate by pollen tube.
(2) Siphonogamy & This type of fertilization occur in higher gymnosperms. Male gametes are non motile transferred to female gamete (egg) by pollen tube. After Pollination male & female gametes are fused & form a diploid zygote.
Life Cycle Of Gymnosperm
Development Of Zygote –
1. Embryo is formed by the development of diploid zygote. After embryo formation ovule is termed as seed. (Seed = Ovule + Embryo)
2. Embryo is enclosed in seed. Seed absorbs water & bursts. Now embryo germinates and forms a new diploid plant.
Note :- Different types of polyembryony are found in gymnosperm i.e. a single seed develops many embryo.
LIFE CYCLE :
Life cycle of Gymnosperm & angiosperm is diplontic because gametophytic generation is short lived. Gametophyte is very reduced & depends on its sporophyte.
SOME IMPORTANT POINTS :
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Evolution of archegonia starts form liverworts. It is well developed in moss. Gymnosperm is last group of it. So in this group it is very reduced. Archegonia of Liverworts – NCC = 4 - 6 VCC = 1 Egg cell = 1 Archegonia of Moss – NCC = 6 - 18 VCC = 1 Egg cell = 1 Archegonia of Pteridophyta – NCC = 1 - 2 VCC = 1 Egg cell = 1 Archegonia of Gymnosperm – NCC = Absent VCC = 1(Short lived) Egg cell = 1 |
2. During evolution Gametophyte becomes reduced & sporophyte becomes well-developed. Gametophyte
Sporophyte Well developed – In Angiosperm |
Gymnosperms are divided into two groups (1) Cycadophyta (Lower Gymnosperm) |
Cycadophyta
(A) The plants of this group are megaphyllous or macrophyllous with circinate vernation
(B) Presence of Ramenta.
(C) Male gamete is motile.
(D) Vascular bundles are hadrocentric in fossil members.
(E) Stem is caudex and wood is mannoxylic.
Cycadophyta is divided into three orders:
(i) Cycadofillicales or Pteridospermae :-
(a) This group includes seed ferns. Many characters of ferns are found in this group like megaphyllous leaves, circinate vernation, ramenta, motile gametes, hadrocentric vascular bundles.
(b) Now this group is completely extinct. eg. Lyginopteris - Fossil plant
(ii) Benettitales :-
(a) It is also a completely extinct group. eg. Williamsonia - fossil plant
Note : Its fossils were discovered by Prof. Birbal sahani
(iii) Cycadales -
(a) Presently living cycadophytes are included in this order.
(b) All the plants of this group are living fossils.
CONIFEROPHYTA
Four orders are included in this group
(i) Ginkgoales :-
(a) It is the oldest order of coniferophyta. Maximum plants of this group are extinct.
(b) Only one plant of Ginkgo biloba is present in India (In Manali). Some plants are also present in China.
Ginkgo biloba -living fossil - It is also known as "Maiden hair tree".
Note : Exceptionally Ginkgo biloba belongs to higher gymnosperm but its male gametes are motile and fertilization by zoodiosiophonogamy.
(ii) Cordaitales :-(a) It is completely extinct group eg. Cordaites
(iii) Coniferales :-
(a) Conifers are included in this group.
(b) It is the largest group of gymnosperm e.g. of Coniferales :-
(a) Pinus (Pines) :-
(b) Cedrus – It is known as deodar.
(c) Taxus – It is known as Yew tree. An anticancer medicine "Taxol" is obtained from its bark.
(d) Taxodium maxicanus – The stem of this plant is thickest in the plant kingdom.
(e) Abies balsamea – A resin "Canada balsam" is obtained from it. It is used to manufacture permanent slides in biology laboratory.
(f) Juniperus virginiana – An oil is obtained from this tree - "Cedar wood oil". It is used as cleansing fluid in biology laboratory. This oil is also used in microscope to increase the resolving power. This oil is used as nail polish remover.
(g) Araucaria species
(h) Sequoia species – The plants in this genus are heavy. Therefore they are called as father of forest.
Sequoia giganteum – It is called Redwood tree or Sherman tree. It is the largest gymnosperm. There is only one tree of this species and that is in California (America)
(i) Metasequoia – It is a living fossil. This plant is present in china valley.
iv) Gnetales -
(a) They are the most advanced gymnosperms.
(b) Exceptionally members of this group have vessels in xylem.
(c) Ovule of this group is bitegmic.
(d) Archegonia is absent in the members of this group. eg.
(1) Gnetum (2) Welwitschia (3) Ephedra – Exceptionally archegonia is present in Ephedra.
Ephedra- This gymnosperm is commonly found in Rajasthan. Ephedra is a medicinal plant. Ephedrine (Medicine) is obtained from it. It is an effective medicine in asthma. Athletes misuse it, so ephedrine is restricted for them.
Cycadofillicales were first seeded plants. First time seed habit was established in this group. In general way seed habit was originated from those pteridophytes which are now extinct. These pteridophytes were ancestor of cycadofillicales. Seed habit was originated from those pteridophytes which were just like Selaginella. Because Selaginella show origin of seed habit. There is mainly two requirements for seed formation (1) Plant should be heterosporus |
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