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Fertilization, Formation and Foetal Membrane in Mammals | Animal Husbandry & Veterinary Science Optional for UPSC PDF Download

 Fertilization

  1. Fertilization Process:

    • The head of the sperm enters the egg, and this joining is called fertilization.
    • After entering, the sperm head rotates and moves toward the center of the egg.
  2. Chromosomal Changes:

    • The nuclei (containing genetic material) of both the egg and sperm come together.
    • The protective membranes around these nuclei disappear, and the genetic material combines to form chromosomes.
  3. Cell Division:

    • The centrosome of the sperm divides into two and positions itself between the newly formed chromosome groups.
    • This marks the start of cell division.
  4. Result of Fertilization:

    • The original number of chromosomes, which is unique to each species, is restored through fertilization.
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Formation of Germ Layers in Mammals

  1. Cleavage Process in Mammals:

    • Cleavage is the process where the fertilized egg (zygote) divides into smaller cells, known as blastomeres.
    • This cell division continues until a certain number of cells are formed, creating a structure similar to the original zygote.
  2. Formation of Blastula:

    • Blastomeres arrange themselves in a circular manner, forming a structure called the blastula.
    • The blastula has a central space known as the blastocoel.
  3. Structure of Blastula:

    • The blastula is a hollow sphere lined externally by blastomeres, which form the trophoblast.
    • Inside the blastula, there is an inner cell mass attached to the trophoblast, and this structure is called a blastocyst.
  4. Development of Germ Layers:

    • The inner mass separates into an outer layer called ectoderm and an inner layer called endoderm.
    • The endoderm forms a roof for the yolk sac, and together, these layers are referred to as the embryonic disc.
  5. Primitive Streak and Mesoderm Formation:

    • A primitive streak appears, and cell proliferation in this area produces cells of the mesoderm.
    • The mesoderm layer differentiates into the notochord and two lateral sheets that form the somatic and splanchnic layers.
  6. Formation of Body Cavity:

    • The somatic mesoderm and ectoderm together create the body wall, known as Somatopleurae.
    • The endoderm and splanchnic mesoderm together form the gut wall, known as Splanchnopleurae.

Fig: Formation of germ layers in mammalsFig: Formation of germ layers in mammals

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What is the structure formed by blastomeres arranging themselves in a circular manner called?
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Foetal Membranes and Placentation

  1. Fetal Membranes and Appendages:

    • The set of structures surrounding and supporting the developing fetus include the yolk sac, chorion, allantois, and amnion.
    • The yolk sac, although not functional in mammals, is important as it's where blood cells first appear.
  2. Amnion's Protective Role:

    • The amnion is like a protective bag that surrounds and suspends the embryo.
    • It keeps the embryo safe from drying out and helps maintain its shape without distortion.
  3. Muscle Fibers and Contractions:

    • Muscles in the mesodermal layer of the embryo develop and contract rhythmically.
    • These contractions gently move the embryo, preventing it from sticking to the surroundings.
  4. Allantois as a Temporary Storage:

    • The allantois serves as a temporary sac for storing the fetus's urine.
  5. Placenta's Vital Role:

    • The chorion, covered in vascular villi, interacts closely with the endometrium (lining of the uterus).
    • This interaction forms the placenta, which is crucial for nutrition, respiration, waste elimination, and protection of the developing embryo.

Fig: Formation of octal membranesFig: Formation of octal membranes

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FAQs on Fertilization, Formation and Foetal Membrane in Mammals - Animal Husbandry & Veterinary Science Optional for UPSC

1. What is fertilization and how does it occur in mammals?
Ans. Fertilization is the process of fusion of male and female gametes, resulting in the formation of a zygote. In mammals, fertilization occurs internally. The male gametes, called sperm, are deposited into the female reproductive tract during sexual intercourse. The sperm then travel through the female reproductive tract and reach the site of fertilization, which is usually the fallopian tubes. Here, a single sperm penetrates the egg cell, forming a zygote.
2. What are germ layers and how are they formed in mammals?
Ans. Germ layers are the three primary layers of cells that form during embryonic development and give rise to different tissues and organs in the body. In mammals, the three germ layers are the ectoderm, mesoderm, and endoderm. The formation of germ layers in mammals occurs through a process called gastrulation. During gastrulation, the cells in the embryo undergo extensive rearrangement and migration. The ectoderm forms the outer layer of the embryo, giving rise to the skin, nervous system, and sensory organs. The mesoderm forms the middle layer, giving rise to muscles, bones, and the circulatory system. The endoderm forms the innermost layer, giving rise to the digestive system, respiratory system, and other internal organs.
3. What are foetal membranes and what is their role in mammals?
Ans. Foetal membranes are specialized structures that surround and protect the developing embryo/fetus in mammals. They provide essential functions for the growth and development of the fetus. The foetal membranes in mammals include the amnion, chorion, yolk sac, and allantois. The amnion is a fluid-filled sac that cushions the fetus and provides protection against mechanical shocks. The chorion is involved in gas exchange between the fetus and the mother's blood. The yolk sac initially provides nutrients to the developing embryo and later contributes to the formation of the digestive system. The allantois stores waste products and facilitates gas exchange.
4. What is placentation and how does it occur in mammals?
Ans. Placentation is the process by which a placenta is formed in mammals. The placenta is an organ that develops during pregnancy and serves as the interface between the mother and the developing fetus, providing nourishment, oxygen, and waste removal. In mammals, placentation occurs through the fusion of the maternal and fetal tissues. The chorion, a foetal membrane, establishes contact with the maternal tissues, forming specialized structures called placental villi. These villi contain blood vessels and allow the exchange of nutrients, gases, and waste products between the mother and the fetus. Placentation is crucial for the survival and development of the fetus throughout gestation.
5. How do fertilization, germ layer formation, foetal membrane development, and placentation contribute to mammalian embryogenesis?
Ans. Fertilization marks the beginning of embryogenesis in mammals. It results in the formation of a zygote, which undergoes cell division and differentiation to form an embryo. Germ layer formation is a critical step in embryogenesis as it gives rise to all the major tissues and organs in the body. The ectoderm, mesoderm, and endoderm differentiate into specific cell types and contribute to the development of various body systems. Foetal membranes play a vital role in the protection and nourishment of the developing embryo/fetus. The amnion provides a cushioning fluid environment, while the chorion, yolk sac, and allantois are involved in gas exchange, nutrient uptake, and waste removal. Placentation is the establishment of the placenta, which ensures the exchange of nutrients, gases, and waste products between the mother and the fetus. It plays a crucial role in supporting the growth and development of the fetus throughout pregnancy.
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